Abstract
Dryland pastoral systems across Africa are routinely characterised as ecologically fragile and economically marginal, yet the herding communities that inhabit them have, over centuries, developed detailed, place-calibrated knowledge of how rainfall pulses translate into forage. In Richtersveld National Park (RNP), South Africa's Northern Cape, the Nama people's annual transhumance between Succulent Karoo uplands and the Orange River riparian zone is one such system. Drawing on stratified vegetation surveys, allometric browse measurements, satellite-derived productivity data, and soil nutrient profiling conducted across five vegetation types between 2006 and 2008, this study examines the ecological mechanisms underlying this pastoral system. Annual plant biomass was most strongly predicted by rainfall (R² = 0.69, 95% CI: 0.56–0.79; F(1,58) = 36.90, p < 0.01), with grazing pressure and landform accounting for far less variance. Shrub biomass in the Succulent Karoo peaked at 962.8 g/m², nearly twelve times the 81.6 g/m² recorded in the Desert Biome. Riparian grass under low-discharge conditions reached 600–700 g/m² during months when all upland vegetation was dormant. MODIS-derived fPAR values tracked ground-measured biomass closely (R² = 0.89). The results provide quantitative evidence that year-round forage availability is maintained under the existing management regime; whether this translates into sustained livestock body condition or reproductive performance under future climate conditions requires dedicated herd-monitoring data that were beyond the scope of this study. Taken together, the results provide a quantitative ecological rationale for practices that policy has historically dismissed as traditional rather than adaptive.
Keywords
References
- Baker, L. E., & Hoffmann, T. M. (2006). Managing variability: Herding strategies in communal rangelands of semi-arid Namaqualand, South Africa. Human Ecology, 34, 765–784. [Google Scholar]
- Fynn, R. W. S. (2012). Functional resource heterogeneity increases livestock and rangeland productivity. Rangeland Ecology and Management, 65, 319–329. [Google Scholar]
- Hempson, G. P., Illius, A. W., Hendricks, H. H., Bond, W. J., & Vetter, S. (2015). Herbivore population regulation and resource heterogeneity in a stochastic environment. Ecology, 96(8), 2170–2180. [Google Scholar]
- Hendricks, H. H., Clark, B., Bond, W. J., Midgley, J. J., & Novellie, P. A. (2005b). Movement response patterns of livestock to rainfall variability in Richtersveld National Park. African Journal of Range and Forage Science, 22(2), 117–125. [Google Scholar]
- Hendricks, H. H., Midgley, J. J., Bond, W. J., & Novellie, P. A. (2004). Why communal pastoralists do what they do in the Richtersveld National Park. African Journal of Range and Forage Science, 21, 29–36. [Google Scholar]
- Hendricks, H. H., Novellie, P. A., Bond, W. J., & Midgley, J. J. (2005a). Plant species richness and composition along livestock grazing intensity gradients in a Namaqualand South Africa protected area. Plant Ecology, 176, 19–33. [Google Scholar]
- Kimiti, K. S., Western, D., Mbau, S. J., & Wasonga, O. V. (2018). Impacts of long-term land-use changes on herd mobility in Amboseli Ecosystem, Kenya. Ecological Processes, 7(4), 1–9. [Google Scholar]
- Konje, M. M. (2021). Spatial-temporal variation in forage resource production in Richtersveld National Park, South Africa [Doctoral thesis, Kenyatta University]. [Google Scholar]
- Konje, M., Muoria, P., & Wabuyele, E. (2021a). Spatial-temporal variation of biomass production by shrubs in the Succulent Karoo, South Africa. Journal of Biodiversity and Environmental Sciences, 18(4), 82–92. [Google Scholar]
- Konje, M., Muoria, P., Wabuyele, E., & Vetter, S. (2021b). Spatio-temporal variation in forage production in a key resource area in the Succulent Karoo rangeland, South Africa. Researchjournali's Journal of Ecology, 8(1), 1–21. [Google Scholar]
- MacKellar, N. C., Hewitson, B. C., & Tadross, M. A. (2007). Namaqualand's climate: Recent historical changes and future scenarios. Journal of Arid Environment, 70(4), 604–614. [Google Scholar]
- Mucina, L., & Rutherford, M. C. (2006). The vegetation of South Africa, Lesotho and Swaziland. South African National Biodiversity Institute. [Google Scholar]
- Naah, J. S. N., & Guuroh, R. G. (2017). Factors influencing local ecological knowledge of forage resources: Ethnobotanical evidence from West Africa's savannas. Journal of Environmental Management, 188, 297–307. [Google Scholar]
- Otieno, J., Wambua, L., & Gacheru, E. (2019). Secondary metabolites and palatability of browse species in semi-arid rangelands. African Journal of Ecology, 57, 420–430. [Google Scholar]
- Samuels, M. I., Allsopp, N., & Hoffman, M. T. (2013). How could herd mobility be used to manage resources and livestock grazing in semi-arid rangeland commons? African Journal of Range and Forage Science, 30(1&2), 85–89. [Google Scholar]
- Samuels, M. I., Allsopp, N., & Hoffman, M. T. (2019). Traditional mobile pastoralism in a contemporary semi-arid rangeland in Namaqualand, South Africa. Rangeland Ecology and Management, 72(10), 195–203. [Google Scholar]
- Smet, M., & Ward, D. (2006). Soil nutrient contents and electrical conductivity affect plant species diversity in semi-arid rangelands in South Africa. Journal of Arid Environments, 67, 54–64. [Google Scholar]
- Young, J. A., Guo, D., Desmet, P. G., & Midgley, G. F. (2016). Biodiversity and climate change: Risk to dwarf succulents in Southern Africa. Journal of Arid Environment, 129(1), 16–27. [Google Scholar]

